Hello,
To clarify re. switch 2,4, 'Always Closed', it's used to show the software that the '12v Digital' used for the switch matrix is okay; if it's absent, the program halts after booting up, showing a message about checking some fuses, the presumed cause.
As to the actual connection, it's made on the 'Coin Door Interface' pcb, in the cabinet. A diode is connected between Column 2 and Row 4, without any 'switch' as such. So booting up a Wpc game without the CDI board connected (to the machine, the door could be missing, it wouldn't matter) will see the warning message appear.
Calimori, the 8-driver board can be configured as split between 6 solenoid drives and 2 switch Columns - even Star Trek Next Gen didn't need both of them. Though I see now that it gave up the usual 'Ticket Opto' as 2,3. Which seems to have been the last of it, 2,3 then became the preferred location for Buy-In button.